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This module introduces the processes involved in maintaining genome stability, causing genome variability and controlling the coding potential of the genome. Mutation, recombination and transposition, and the interplay between them, are examined as causes of genome instability. The two main themes of the module are the impact of genome instability/change upon gene expression, and its control. An introduction to bioinformatics and sequence analysis with the use of sequence databases and analysis tools permits the analysis of gene/genome variability with an introduction to the importance of bioscience research, including molecular diagnostics and drug development.